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HK-2 cells were maintained in Dulbecco’s modified Eagle’s medium/nutrient mixture F-12 (DMEM/F12, Gibco, USA), whereas HEK293 cells were maintained in DMEM basic medium (Gibco, USA), both supplemented with 10% fetal bovine serum (FBS, Clark USA) and 1% penicillin/streptomycin antibiotics (NCM, China) and incubated at 37 °C with 5% CO 2

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Introduction Gallbladder cancer (GBC) is an aggressive malignancy associated with a dismal prognosis, frequently diagnosed at advanced stages with scarce therapeutic options